Harold W. Keller

Resident Research Associate


Ph.D. in Botany, University of Iowa

M.A. in Botany, University of Kansas

B.A. in Biology, Kansas Wesleyan University

Post Doctoral Fellow in Botany, University of Florida

Harold is a native Kansan, who grew up on the edge of the Flint Hills at Peabody. He joined BRIT as a Research Associate in 1990 and has published more than 40 book reviews, 60 journal papers, 116 abstracts, and one book entitled Myxomycetes of Ohio: Their Systematics, Biology, and Use in Teaching. He has given presentations at professional meetings such as The International Mycological Congress, International Congress on the Systematics and Ecology of Myxomycetes. (ICSEM 2, 3, 4, 5, 6, 7) and he delivered the invited Plenary Address at ICSEM2 held at Madrid, Spain, at the Mycological Society of America (MSA), and Association of Southeastern Biologists (ASB), and North American Mycological Association (NAMA) annual meetings. Numerous myxomycete workshops titled “The Wonderful World of Myxomycetes” and “Myxomycete Systematics Workshop” were presented at ICSEM, MSA, and NAMA meetings. Courses taught include Principles of Biology: Organismic and Ecology, Microbiology and Human Diseases, Biology of Lower Plants, Plant Biology, Biology of Economic Plants, Medical Parasitology and Mycology, Biological Literature, History of Biology, and Biology of Slime Molds at different universities, most recently the University of Central Missouri (UCMO).

His research on the biology, ecology, systematics, and tree canopy biodiversity of myxomycetes and fungi was supported financially by grants from the National Science Foundation, National Geographic Society, Discover Life in America, and the Missouri Department of Natural Resources. His students at UCMO used the Double Rope Climbing Method to access, survey, and inventory the tree canopy for cryptogams in the Great Smoky Mountains National Park and the Daniel Boone National Forest in Kentucky and this activity was highlighted in a recent issue of the Journal of the Botanical Research Institute of Texas (JBRIT). Work presently with the BRIT Press involves publishing earlier research on the anatomy of the genus Echinacea, especially the epidermal cells of the ray flowers. He currently serves on the Editorial Board for The BRIT Press and as a Contributing Editor for the new journal FUNGI.

Select Publications:

Keller HW, SE Everhart, CM Kilgore. 2021. Myxomycetes: Basic Biology, Life Cycles, Genetics and Reproduction. Pp. 1–45 in: Myxomycetes: Biology, Systematics, Biogeography and Ecology, 2nd Edition, edited by C Rojas, S Stephenson. Elsevier An Imprint of Academic Press, Atlanta, GA.

Keller HW. 2021. Morel mushrooms, macrofungi, corticolous myxomycetes, and American elm trees. FUNGI 14:23-34.

Keller HW. 2020. The Cross Shot: Grave Peak Lookout, Powell Ranger District, Idaho. Lookout Network 30(3-4):23.

Keller HW, RL Price, BG Stone, ED Forrester. 2020. Arcyria versicolor of western mountains, U.S.A. (Myxomycetes: Trichiales: Trichiaceae): A morphological and taxonomic study with observations of nivicolous species. Journal of the Botanical Research Institute of Texas 14(2):435-459. doi.org/10.17348/jbrit.v14.i2.1021

Lopes-Andrade C, ML Ferro, HW Keller. 2020. A new species of Cis Latrielle (Coleoptera: Ciidae) from the USA, with comments on the use by Ciidae of Stereaceae Fungi (Basidiomycota: Agaricomycetes: Russulales) as hosts. The Coleopterists Bulletin 74(1):93–100. doi.org/10.1649/0010-065X-74.1.93

O’Kennon, B, HW Keller, BA Bunyard. 2020. Observation of the stinkhorn Lysurus mokusin (Phallales: Phallaceae), and other fungi found on the BRIT campus in TexasFUNGI 13(3):41–48.

Perry BA, HW Keller, ED Forrester, BG Stone. 2020. A new corticolous species of Mycena sect. Viscipelles (Basidiomycota: Agaricales) from the bark of a living American Elm tree in Texas, U.S.A. Journal of the Botanical Research Institute of Texas 14(2):167-185. doi.org/10.17348/jbrit.v14.i2.1000

Keller HW. 2019. Student team-based tree canopy biodiversity in Great Smoky Mountains National ParkPlant Sci. Bull. 65(1): 28–37.

Keller HW, VM Marshall. 2019. A new corticolous myxomycete species (Licea: Liceaceae: Liceales) and crystals on American elm tree bark in Texas, U.S.A. J. Bot. Res. Inst. Texas 13(2):367‒386. 

Keller HW, KL Snell. 2018. Hunting and collecting Myxomycetes at night with a flashlight. FUNGI 11(2):43–44. www.fungimag.com/summer-2018-articles/Flashlight%20LR.pdf

O’Kennon B, D Benjamin, HW Keller. 2018. Xylobolus frustulatus (Stereaceae): Developmental observations, morphology, and ecology. FUNGI 10(4):16–21. https://bit.ly/2JiUdR0

O’Kennon, B, HW Keller, BA Bunyard. 2020. Observation of the stinkhorn Lysurus mokusin (Phallales: Phallaceae), and other fungi found on the BRIT campus in TexasFUNGI 13(3):41–48.

Keller, HW, KM Barfield. 2017. The Great Smoky Mountains National Park: The People’s ParkFUNGI. 10(2): 44–64.

Keller HW, SE Everhart, CM Kilgore. 2017. The Myxomycetes: Basic biology, life cycles, genetics and reproduction. Chapter 1, pp. 1–40, in Myxomycetes: Biology, systematics, biogeography and ecology. Edited by S Stephenson, C Rojas. Elsevier, Atlanta, GA.

Keller HW, B O’Kennon, G Gunn. 2016. World record myxomycete Fuligo septica fruiting body (aethalium). FUNGI 9(2):6–11.

Keller, HW. 2015. More than 50 years with Myxomycetes (plasmodial slime molds): Highlights and review. Taxonomy and Ecology of Myxomycetes. Higher Education Progress. 9-28, Beijing, China. Keynote address at the Eigth International Congress on Systematics and Ecology of Myxomycetes held in Changchun, China (People’s Government of Jilin Province).

Keller, HW. 2014. The genus Echinacea (Asteraceae): Floral, stem, and petiole morphologyJournal of the Botanical Research Institute of Texas. 8:87–126.


Contact Details

Phone: (817) 460-6695

Email: hkeller@brit.org

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